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    ???????????? ????????? ?????? ??????? ????????????? ??, ????????? ??????? (????) ?? ?? ??? ???? ???, ????????? ?????? ????????? ?? ??????? ?????? ??????? ?????? ?????? ?? ?????? ????????? ????????? ?????? ??? ?????. ??? ????, ????????? ??????? ???? ???????????? ??? ???? ?? ???? ?????? ????? ?????? ?????? ???????? ??? ?????.

    ??? ???????, ????? "?????? ??? ???, ????????? ??? ???" ???? ??????? ?????, ?? "??????" ?????? ???? ???? ??????? ?????? ????. ????????? ??????? ?? ?? ?????????? ????????? ???? ???. ?? ????? ???????? ??? ???? ????????? ????? ????????? ??????, ???????? ???????????? ???????? ??????????? ?????? ??, ?? ????????? ???????????? ???????? ?????????? ????? ??? ???? ??? ????????? ???????? ?????? ????.

    ?? ??????? ???? ????, ????? ???????? ?????? ??????????? ???????? ???? ??????? ??????? ????? ?????? ?? ????? ????????????? ???????? ?????????? ???????? ????: ????????? ?? ???????????????? ?????? ????-???????? ????? ??????? PI ?-????? VPS?? ?????? ?????? ????.

    ?????????? ??? ????? ??? ??? ?? ????-???????? ?????????, ???????? (????????? ???), ????????? ?????????? ?????? ??? ??? ????. ???????? ?? ????????? ?? ?? ??????? ????????? ????????? ???, ?? ????????? ??????? ???????????????? ????? ??? ?????? ??????? ????? ???????????? ???? ???????? ?????????? ???? ??? ????????? ???????? ??? ???? ???.

    ??????????? ?????????? ??? ????? ??? ??? ?? VPS34 ?? ???????????????????? 3-????? (PI3K) ?? ?? ?????? ?????? ???, ???????? VPS34 ?? ?????? ???? ????, PI3P ?? ??????? ??? ??? ???? ??? ????????? ???????????? ????????? ??????? ???????? ???????????? ??? ????? ????. ?????, ??????? ???? ?????? ?????? ??? ???????? ?? ????????? ???? ?????????? ?????? ????? ???? ????. ????????????, ?????? ?????? ?? ?????? ???? ?? X-?????? ????????????? ???????? (XLMTM) ???????? ???? ????????? ??????? ?????????? ?????????? ?????? ????? ???.

    ?? ?????????????? ????????? ????????? ???????? ?? ???? ?? ??????? ????? ??? ???? ??? ????? ???????????????? ?????????? ???????? ??? ????????? ????????? ??????????? ???? ?????? ?????.

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    ?? ????? ?????????? ???? ????????? ????-???????? ?????????, ???????? ?????? ??????? (??????) ?????? ????? ????, ?? ????? ?????????????? ????????? ?? ?? ????????? ???. ??????? ????? ?? ?????????? ??????? ???????? ??????? ????????? ????????? ???????????? ???????? ???? ???? ??? ??????????? ????????????? ?????? ??????? ??? ????.

    ???????, ??????????? ???????? ???????? ???????? ??? ????? ???? ??? ?? MSB ?? ???? ?????? ?????? ????-???????? ??????????? ????????? ????????? ???????????? ????????? ???????? ?????????? ????? ??? ????. ???? MSB PI3K kinase VPS34 ?? ?????? ???? ????, PI3P ?? ??????? ??? ??? ???? ??? ???????????? ??? ????? ????, ???????? ????????? ???????????? ??????? ?????? ????. ????????????, X-?????? ?????????????? ???????? (XLMTM) ?????, MSB VPS34 ?? kinase ?????????? ???? ????, PI3P ?? ??????????? ??????? ??????? ??? ???????? ??????????? ??????????? ??? ????.